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Selfish learners can converge to persistent cooperation in social dilemmas, but only if their learning rates are heterogeneous.
A "marginal feedback" payoff structure, rewarding pivotal opinions, enables layered populations to evolve collective intelligence for complex non-linear tasks, mirroring multi-layer perceptrons.
Higher pathogen prevalence actually slows down antigenic evolution, overturning the common assumption that larger infected host volumes inevitably accelerate adaptive escape.